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Evolution—History of life.
Evolution is the process by which new species are formed from pre-existing one of a long period of time. It is believed that life exists only on earth of all planets.
Specific objectives
The learner should be able to
Theories for origin of the earth
3. It was the design of a creator.
Theories for origin of life on earth.
1. Special creation; life was created by a supernatural being at a particular time. Genesis 1; 1- 26
2. Spontaneous generation; life arose from non- living matter on numerous occasions.
3.Steady-state. Life has no origin
4. Cosmozoan; life arrived on this planet from elsewhere.
5Biochemical evolution; life arose according to chemical and physical laws.
Modern view.
Modern geologists believe the earth is over billion years old. They speculate that the mountains deserts and oceans of today formed from slow, gradual but continuous process of erosion and uplifting. The about 3.5 billion year ago, life began From simple unicellular organism, new life forms arose and changed in response to environmental pressures producing the past and present biodiversity [evolution]
Theories of evolution.
Evolution is an overall gradual development that is both ordered and sequential. In terms of living organisms it may be defined as; the development of a differentiated organisms from pre-existing less differentiated organism over the course of life.
Lamarckian theory.
The French biologist Lamarckian proposed, in 1805 a hypothesis to account for the mechanism of evolution based on two conditions; The use and disuse of parts and inheritance of acquired characteristics. Changes in the environment may lead to changed patterns of behavior which necessitates new increase use or disuse of certain organisms/or structures. Extensive use would lead to the increased size and or efficiency whilst disuse would lead to degeneracy and a trophy. These traits acquired during the lifetime of individuals were believed to be heritable and thus transmitted to offspring.
According to Lamarckism, as the theory came to be to known, the long neck and legs of the modern giraffe were the results of the generation of short-necked and legged giraffe ancestor feeding on leaves at progressively higher levels trees. The slightly longer necks and legs produced in each generation were passed on the subsequent generation until the size of the present-day giraffe was reached.
Darwin, Wallace and the origin of species by natural selection
Darwin and Wallace proposed that natural selection is the mechanism by which new species arise from pre-existing species. This hypothesis/theory is based on three observations and two deductions which may be summarized as follows:
Observation 1;
Individuals within a population have a great reproduction potential.eg American oyster produces 10 eggs per season.
Observation 2;
he number of individual in a population remain approximately constant
Deduction 1;
Many individuals fail to survive or reproduce. There is a ‘struggle for existence’ with the population
Observation 3.
Variations exist within all populations.
Deduction 2:
In the ‘struggle for existence,’ those individuals showing variation best adapted to their environment have a ‘reproductive advantage’ and produce more offspring than less adapted organism.
Natural Selection
This is a natural mechanism by which those organisms which appear physically, physiologically and behaviorally better adapted to the environment survive and reproduce. Those organisms not so well adapted fail to reproduce or die. The former organisms pass on their successful characteristics to the next generation.
Variation.
The term variation describes the differences in characteristics shown by organism belonging to the same natural population or species.
Types of variation
Variation is said to be continuous when there is a gradual change of character from one individual to another e.g. skin color, length of leaves, height of individuals. i.e. it is quantitative.
2. Discontinuous variation
In discontinuous variation, there is a clear – cut difference between the characteristics e.g. blood group tongue- rolling, sex etc.
Variation and selection
Selection is the process by which those organisms which appear physically, physiologically and behaviorally well adapted to the environment survive and reproduce. They pass on their successful characteristic to the next generation while the less adapted die.
Therefore, selection can be seen to operate through the process of differential mortality and differential reproduction potential. The selection has an adaptive significance in perpetuating those organisms most likely to ensure the survival of the species and depends upon the existence of phenotypic variation with the population as food availability in animals and light in plants. This produces competition for resources between members of the population. Those organisms exhibiting characteristics which give them a competitive advantage will obtain the resources, survive and produce organism without those characteristics are at a disadvantage and may die before reproducing.
Both environmental limiting factors and population size operate together to produce a selective pressure which can vary in intensity and from time to time and from place to place.
Types of selection.
Three types of natural selection
1.directional selection
2.stabilising selection
3.disruptive selection
Directional selection
occurs when an extreme phenotype is favored. Such a shift occurs when a population is adapting to a changing environment. E.g.,
Stabilizing selection
occurs when an intermediate phenotype is favored. It can improve the adaptation of a population for those aspects of the environment that remain constant. With stabilizing selection extreme phenotypes are selected against and individuals near the average are favored e.g. birth weight in human infants. Babies heavier or lighter than 3.6 kg in London are at a selective disadvantage and have a slightly increased rate of mortality.
Disruptive selection
Here two or more extreme phenotypes are favored over any intermediate phenotype. Fluctuating conditions in an environment, say associated with season and climate may favor the presence of more than one phenotype within a population. Selection pressure acting from within the population as a result of increased competition may push the phenotype away from the population mean towards the extremes of the population. This can split a population into two subpopulations and can result in different phenotypes with a population phenomenon known as polymorphism.
Graphical illustration of the types of selection
Polymorphism
This is the existence or more forms of the same population and can be applied to biochemical morphological and behavioral characteristics e.g., land snail cepaea nemoralis, the shell species may be yellow, brown or various shades
Types of polymorphism.
This arises when different forms or morphs, exist in a population undergoing a strong selection pressure. The frequency of the phenotypic appearance of each form is determined by the intensity of selection pressure, e.g., melanic and non-melanic form of the peppered moth. Transient polymorphism usually applies in situation where one form is gradually replaced by another
2. Balanced polymorphism.
This occurs when different forms co-exist in the same population in a stable environment. E.g.
Causes of variation
Variations are caused by both genetical and environment differences. From the evolution point of view genetic variations are more important because they can be transmitted from parents to off springs.
The causes of genetic variations
– orientation on the equator in metaphase I
During metaphase I of the first meiotic division homologous chromosomes came together in pairs and subsequently segregate into daughter cells independently of each other. The result of this independent assortment is the production of a wide variety of different gametes depending on which particular chromosome ends up with one another in each cell.
– Crossing over
In prophase of the first meiotic division, homologous chromosomes came together and make intimate contact with each other. Chromatid of homologous chromosome may break and rejoin at any place called chiasmata.
Fertilization
Union of gametes at fertilization results in alleles present in one gamete being united with alleles in another. If a population consist of large number of outbreeding individual, the amount of variation that may result from this is again virtually infinite.
Despite the tremendous amount of variation these three processes may generate, they play only a limited role in evolution. The reason is that although they may establish a new combination of alleles in one generation, they do not generate a long-lasting variation of a novel kind.
Mutation
This is a change in the amount or structure of DNA of an organism. This produces a change in the genotype which may be inherited by cells derived by mitosis or meiosis. Individuals showing the new characteristics are referred to as mutants. Mutants arise spontaneously and in no sense ‘directed’ by the environment, although the environment greatly influence the mutation rate.
Factors or agents that speed up or lead to mutations are called mutagens include; Gamma rays, ultraviolet and a number of chemicals e.g., mustard gas, colchicine in plants
Effect of mutation
2. The vast majority of mutation confers disadvantages on organisms that inherit them.
Causes of mutation
Ultraviolet light
Chemicals
Freezing of cells
Types of mutation.
This may be the result of changes in the number or structure of chromosomes
This is usually due to errors occurring during meiosis and mitosis. The changes may involve the loss or gain of the single chromosome a condition called Aneuploidy or the increase in an entire haploid set of chromosomes a condition called euploidy[polyploidy]
Aneuploidy
In this condition half of the daughter cell produced have an extra chromosome [n + 1], [2n + 1] and so on, while the other half have a chromosome missing [n- 1], [2n- 1] and so on.
An aneuploidy can arise from the failure of a pair or pairs of homologous chromosome to separate during anaphase 1 of meiosis. If this occurs both sets of chromosomes pass to the same pole of the cell and separation of homologous chromosomes during anaphase II may lead to information of gametes containing either one or more chromosome too many or too few. This is known as non-disjunction.
One of the commonest form of chromosomal mutation in human resulting from non-disjunction of the G 2, chromosome is Dawn’s syndrome [2n +1] characterized by mental retardation, reduced resistance to disease, congenital heart abnormalities a short stocky body and thick neck and the characteristic fold of skin over the inner corner of the eyes. Down’s syndrome and other related chromosomal abnormalities occur more frequently in children
born to older women. Klinefelter’s syndrome is due to non-disjunction is sex chromosomes, which may result into individuals who have a genetic constitution, XXY, XXXY, or XXXXY. These individuals are phenotypically male but have small testes and no sperm in the ejaculate. There may be abnormal breast development and the body proportions are generally female. Y is the cause of maleness.
Turner syndrome [2n- 1]: individual inherit a single X-chromosome (XO). Individuals with this condition often do not survive pregnancy and are aborted. Those that do are phenotypically female, but small in stature and sexually immature. Despite having a single X chromosome, like males, they are female, indicating again that the Y chromosome is the cause of maleness
Polyploidy
This occur when there is an increase in the entire haploid sets of chromosomes; i.e., 3n triploid], 4n [tetraploid]
Polyploidy is rare in animals but common in plants. Polyploidy is often associated with advantageous characteristics such as increased sized and greater, the hardness of seeds, though such advantages are sometimes offset by reduced fertility. Polyploidy is sometimes induced by colchicine an alkaloid substance extracted from the crocus colchicum
Structural change in chromosomes
These include loss, multiplication or changes in the sequence of bases on a chromosome
Gene mutation
Gene mutation arises as a result of a chemical change in an individual gene and it ‘s thought to be very important in generating evolutionary change. An alternation in a sequence of nucleotide in a gene may change the order of amino acids making proteins. This may affect the fitness of the organism. It includes.
Somatic mutation.
Here mutation in non-reproductive cells of an organism. The resulting genetic change will present in all descended cells from the original mutant cell and may have a profound effect on individuals.
However, as the genetic change is only in non-reproductive cells, it cannot be transmitted to feature generation.
Gene pool
This Is the total variety of genes and alleles present in asexually reproducing population, and in any given population. The composition of gene pool may be constantly changing from generation to generation as a result of natural selection. Populations undergoing evolutionary change have continuously changing gene pools.
Allele frequency
This the fraction of organisms in a population carrying a particular allele.
Genotype frequency
This is the fraction of organisms in a population carrying a particular genotype. The frequency of dominants and recessive allele in a population will remain constant from generation to generation provided the following conditions exist.
Any change in allele or genotype frequencies must, therefore, result from the alteration of one or more of the conditions above. These are factors that significant in producing evolutionary a
Factors producing a change of genotype are allele in the population.
non- random breeding
Mating in most natural populations is nonrandom. Sexual selection occurs whenever the presence of one or more inherited characteristics that increases the likelihood of bringing about successful fertilization of gametes of some organisms and not in other. There are many structural and behavioral mechanism in both plants and animals which prevent mating from being random, e.g. flowers possessing increased size of petals and amount of nectar are likely to attract more insects and increase the likelihood of pollination. Thus, sexual selection, as a mechanism of non- random mating ensures that certain individuals within the population have an increased reproductive potential so that their alleles are more likely to be passed to the next generation.
Genetic drift.
This refers to the fact that variation in gene frequencies with populations can occur by chance rather than by natural selection. E.g., chance events such as premature accidental death prior to mating of an organism in a small population which is the sole possessor of a particular allele would result in the elimination of that allele from the population.
Genetic load
This is the existence within a population of disadvantageous alleles in heterozygous genotype e.g. sickle cell trait in region where malaria is endemic.
Gene flow.
It’s the movement of alleles from one population to another as a result of interbreeding between members of the two populations.
Heterozygotes as a reservoir of genetic variation (the Hardy-Weinberg principle)
For a particular character in a population, the dominant form expresses itself more often than a recessive form, for example normal skin color is more common than albino. In a large population, the proportion of dominant alleles and recessive alleles of a particular gene remain constant. It is not altered by interbreeding. This constancy is known as the Hardy-Weinberg principle is expressed by a mathematical law
P2 +2pq+ q2 =1
Where p = frequency of allele for dominant character
q = frequency of allele for recessive character
The formula can be used to calculate the frequency of any allele in the population. For example, imagine that a particular metal defect is the result of a recessive allele. If the number of babies born with the defect is 1 in 20000, the frequency of the allele can be calculated as follow:
The defect will only express itself in individuals who is homozygous recessive. Therefore, the frequency of these individuals (q2) = 1/20000 = 0.00005
The frequency of the allele q = √0.00005 = 0.007
Since P + q = 1
The frequency p of the dominant allele = 1- 0.007 = 0. 997
From the Hardy-Weinberg formula, the frequency of heterozygotes is 2pq
i.e., 2 x 0.997 x 0.007 = 0.014
in other words, 14 in 1000 or 280 in 20000 are carriers (heterozygotes) of allele.
This means that in a population of 20000 individuals, one individual will suffer the defect and about 280 will carry the allele. The heterozygotes are acting as a reservoir of the allele, maintain it in the gene pool. As these heterozygotes are normal, they are not specifically selected against, and so the allele remains. Even if the defective individuals are selectively removed, the frequency of the allele will hardly be affected. In our population of 20000, there is one individual who has two recessive alleles and 280 with one recessive allele- a total of 282. The removal of the defective individual will reduce the number of alleles in the population by just 2, to 282. Even with the removal of the defective individual, it would take thousands of years just to halve the allele’s frequency.
Occasionally, as the sickle cell anemia the heterozygotes individual have a selective advantage. This is known as heterozygote superiority.
Conditions that allow the Hardy-Weinberg principle to be true
Speciation
Speciation means the development of different genetic traits in an isolated subpopulation leading to a species distinctly different from the original parent population. Or speciation is the process by which one or more species arise from previously existing species.
A species is a group of organisms that are potentially able to breed amongst themselves but not any other species.
A single specie may give rise to new species [intraspecific speciation] or as is common in many flowering plants two different species may give rise to a new species [(interspecific hybridization). If intraspecific speciation occurs whilst the population are separated it is termed allopatric speciation. E.g. Galapagos island. If the process occurs whilst the population are occupying the same geographical area it is called sympatric speciation
Intraspecific speciation will occur when gene flow with the population is interrupted and each subpopulation is genetically isolated. Then change in allele and genotype frequencies within each subpopulation as a result of natural selection on the range of phenotype produced by mutation and sex recombination lead to the formation of race and subspecies. If genetics isolation persists over a long period of time the subspecies may form new species
Speciation will only occur as a result of the formation of barriers which leads to reproductive isolation between members of the population.
Isolation mechanisms
An isolating mechanism is a means of producing and maintaining isolation within a population. This can be brought about by mechanisms acting before or after fertilization.
Allopatric speciation.
This is characterized by an occurrence at some stage of spatial separation. Geographical barriers such as mountains range, seas, river or habitant preference may produce a barrier to gene flow because of spatial separation. This inability of an organism or their gametes to meet leads to reproductive isolation. Adaption to new conditions leads to a change in alleles and genotype frequencies. Prolonged separation of the population may result in them becoming genetically isolated even if brought together. In this way new species arise.
This is the speciation that occur within the population in the same geographical areas where reproduction isolation may result from structural, physiological behavior of individuals within a population, e.g. polyploidy in plants.
Artificial selection:
This when breeders’ animals and plant select individuals with the characters that are wanted and allow them to interbreed, individuals lacking the desirable qualities are prevented from breeding. By rigorous selection over many generations special breeds or varieties may be developed for a particular purpose.
Animals that have been subjected to artificial selection include.
-cow for beef and milk
-Sheep for wool and meat
-horse for racing and holing
-pig for bacon and lard production
-dog for beauty
Among plants crops such as wheat, barley and potatoes, have been bred for higher yield, greater resistance to disease and drought.
Inbreeding.
This is the crossing of closely related individuals. Inbreeding leads to a loss of fitness known as in breeding depression. This is because an individual produced as a result of the crossing of the close relatives is more likely to have two copies of harmful or ever lethal recessive alleles.
Hybrid.
Is the result of a cross between individuals belonging to two different varieties [out breeding] such individuals show hybrid vigor. This is because hybrid tends to be heterozygous at many of their loci. Any harmful allele, therefore, has its effect masked by healthy ones.
The green revolution
This is the production of new varieties of the world’s major food crop such as rice, wheat, maize and barley by agriculturalists in the recent past.
In general, the new varieties display some or all the following advantages over the older one.
The disadvantage of a green revolution.
New varieties required a high level of fertilizer which is expensive and not always available in developing countries introduction of these species in the developing world concentrates wealth in the hands of the minorly of farms able to afford artificial fertilizer.
EVIDENCE OF EVOLUTION.
The earth is believed to have been composed of one continent from which different day present continents involve apart. For instance, the following provides evidence that the eastern coast of South America was once in contact with the western coast of Africa.
The distribution of animals and plant on different continent provide due to the possibility of evolution as described in the following;
Example
Comparative study of the anatomy of groups of animals and plants[morphology] released that certain structural features are basically similar. For example, the basic structure of all flower consists of sepals, petals, stamen, stigma, style and ovary, yet the size number of parts and specific structures are different for each individual. Similarly, the limb- a bone pattern of all tetrapod from amphibian to mammal has the same structural plan; called the pentadactyl limb. These common basic structures on organisms suggest common ancestry. In each case, structure are modified for a particular function in a particular environment. Some homologous structures that fail to develop full functions and/or structure are referred to as vestigial organs.
3. Homologous structure
These are organs that have a similar basic structure, similar topographic relationship as structure in other species the same histological appearance, similar embryonic development but showing adaptation to different environmental condition and modes of life. E.g., wings of bird and arms of man, Halters of Diptera [housefly] and hind wing many flies.
4. Adaptive radiation
This is a term used to describe the differentiation of homologous structure to perform a variety of function e.g., mouth parts of insects consist of the same basic structure but maxilla in butterfly is modified for suckling and manipulating food in grasshopper;
5. Divergent evolution.
This is the evolution that leads to modification of the basic pattern of homologous structures to enable the descendants to fill a wide variety of ecological niches. Divergent evolution leads to adaptive radiation.
6. Analogous structure
These are structure of organism bearing no close phylogenetic link but showing adaptation to perform the same function e.g.,
-wings of insects and bats
-the jointed legs of insects and vertebrates
Convergent evolution
This is the evolution where structure believed to have different ancestry origin are modified to perform the same basic function. E.g. wings of bird and insects
Vestigial structures.
These are structures that have been greatly reduced or lost from organisms. For example, the third digit in birds’ wings is very much reduced while the fourth and fifth are missing. The existence of vestigial structure has been used as evidence for evolution. It is thought that vestigial structure performed a function in the ancestor but has since been reduced to such an extent that they are lost or greatly changed their original function.
Pentadactyl limb.
It is so-called because typically has five digits is formed in all four classes of terrestrial vertebrate [amphibian, reptile, birds, and mammals] some of the limbs bone can even be traced back the fins of certain fossil fishes from which the first amphibians are thought to have evolved.
Through divergent evolution the pentadactyl limb have become adapted for different function in some species some or all the foes or finger have been lost eg
Seal – for swimming
Human – for manipulation / grasping
Horse – for running
Mole – for digging
Bat or bird – for flight
Similarities in embryonic development stages are thought to have evolutionary significance
Species believed to have a close evolutionary past have been found to show similar embryonic stages. For example, the presence of branchial groves and segmented myotomes in the human embryo is suggestive of a fish ancestry.
Paleontology
This is the study of animals and plants of the past as seen in the fossil records. Fossils are any form of preserved remains through to be derived from a living organisms. They include entire organism, hard skeletal structure, molds and casts, petrifactions, impression, imprints, and coprolites [fossilized fecal pellets]
Embryology
Similarities in embryonic development stages is though to have evolutionary significance
Species believed to have a close evolutionary past have been found to show similar embryonic stages. For example, presence of branchial groves and segmented myotomes in human embryo is suggestive of a fish ancestry.
Paleontology
This is the study of animals and plants of the past as seen in the fossil records. Fossils are any form of preserved remains through to be derived from living organism. They include entire organism, hard skeletal structure, moulds and casts, petrifactions, impression , imprints and coprolites [fossilized fecal pellets]
Ecological consideration also fit in with the fossil evidence. e.g. Plants appeared on land before animals, and insects appeared before insect pollinated flowers
Taxonomy
This is the study of principals, rules and methods of classification. The system of classification proposed by Linnaeus before the time of Darwin and Wallace, based on phylogenetic similarity and differences between organism may suggest existence of evolutionary process. There are similarities and differences between organism may be explained by organism with each taxonomic group to particular environmental conditions over a time.
Numerical taxonomist, working mainly from comparative phenotypic characters have found it possible to construct phenetic classification system which in consistent, to the extent of present knowledge with the concept of evolution organism that have close evolutionary relationship share very many similar characteristics.
Assignment
ASSIGNMENT : Evolution: History of life Assignment MARKS : 10 DURATION : 2 weeks, 1 day